How they interact with other weak interacting particles.
They "carry" lepton familiy number, which is something that must be conserved in an interaction.
for example:
allowed:
[tex]\overline{\nu} _{\mu} + u \rightarrow \mu ^+ + d[/tex]
Not allowed:
[tex]\nu _{\mu} + u \rightarrow \mu ^ + + d[/tex]
where u and d are up- and down- quarks.
in the first reaction, initial muon-family number is -1, and final muon-family number is -1 (since antiparticles has -1, and particles has +1, positive charged leptons are defined to be antiparticles, and negative charged leptons is then particles.)
in the secon reaction, initial muon-family number is +1, and final muon-family number is -1