I notice when lions mate that it is the lioness that makes the most noise. Yes the male does make a deep growl when he grabs the lioness by the neck but the lioness makes a constant high pitched sound that sometimes sounds like a meow. A good example starts at 41:38 in the Lion Brotherhood documentary. There Banda makes a deep growl and the lioness makes a lot of noise. But why does the lioness make the most noise when lions mate. Is it like her way of saying "Ouch, stop doing that"?