The jolt experienced when accelerating in 1st gear is primarily due to the significant difference in gear ratios compared to higher gears. In 5th gear, even without air resistance, the acceleration would not produce the same jolt because the engine operates more efficiently at higher speeds. The discussion emphasizes that gear ratios play a crucial role in how power is transmitted to the wheels. Understanding these mechanics can enhance driving performance and vehicle handling. Overall, the sensation of acceleration is closely tied to the gear in use and the engine's efficiency.