In a sealed elevator accelerating upwards, air molecules do not accelerate uniformly with the elevator due to their inertia. While the elevator and any object in direct contact with it experience the same acceleration, the air inside may create a pressure differential, leading to a sensation of air moving downward. This effect is influenced by gravitational forces when the elevator is on Earth, causing air to distribute unevenly. The discussion highlights that measurement devices can detect subtle differences in air pressure caused by acceleration. Overall, the behavior of air molecules in an accelerating elevator is complex and influenced by both inertia and gravity.