The equation cos²α + cos²β + cos²γ = 1 is significant in crystallography as it relates to the projection of a unit vector in three-dimensional space. When angles α, β, and γ are defined with respect to the x, y, and z axes, this equation holds true, reflecting the Pythagorean theorem in three dimensions. The components of the vector correspond to the cosine of these angles, confirming that the sum of their squares equals one. In two dimensions, a similar relationship emerges, where cos²α + cos²β simplifies to 1 due to the complementary nature of the angles. This illustrates the foundational geometric principles underlying crystallographic structures.