Neurologically, the term reflex is much more specific than the more common usage, which is fine for this discussion.
Reflexes in sports etc. are much slower than many of the more well defined reflexes described in neurobiology (see below) and involve a lot of learned components.
I knew a guy on a softball team I was on who played third base and had incredibly fast reflexes (for catching a ball that get to him rapidly after being hit). My hypothesis would be that he had an innate (somewhat hardwired) reflex to move his arm/hand up to block a rapidly approaching object (perhaps a protection reflex, probably through the
optic tectum) which was linked to learned positioning his hand/glove for catching the ball (probably through the
cerebellum). But who knows. We never stuck him with electrodes while he was doing this.
I would assume most sports reflexes would be at least his complicated, probably more so.Neurologically speaking, reflexes are simple movements or responses triggered but simple stimuli, which usually occur rapidly.
Rapid reflexes involve few neural elements and therefore cross few synapses (communication points between neurons) which is usually where most of the time delay occurs. They are often defined as not involving the brain, but there are many exceptions. They are (mostly) hardwired, not learned.
The classic reflex people learn about is the patellar reflex:
The patellar tendon is stretched (clinically triggered by a gentle tap on the tendon below the kneecap with a rubber hammer).
This activates stretch sensitive sensory neurons which send impulses to the spinal cord (indicating the knee is being bent in an unexpected way).
The sensory neuron endings end on and activate motor neurons that in turn signal the quadriceps thigh muscle to contract, which results in kicking out the lower leg.
Normally, this would be involved in feedback for tuning muscle output to maintain proper leg position under differing loads.
This only involves 2 neurons and 2 synapse (both chemical). Chemical synaptic delays are on the order of 2 ms. Action potentials, which transmit the signals over long distances along axons, go faster in larger and more myelinated axons for
somewhat complex reasons (see myelin and saltatory conduction and cable theory).
My favorite (because I have worked on it) reflex is the
C-start reflex (in fish and amphibeans) which is an escape response to vibrational or contact stimuli. It involves sensory neurons projecting to the hindbrain and stimulating the Mauthner neurons (among others in the area). The Mauthner neuron has the largest axon in the body and therefore has the fastest axon conduction velocity. The Mauthner neurons strongly stimulates motor neurons on the opposite side of the body to fire, causing muscles on the side of the body opposite to the stimulus to contract, moving the animal away from the stimulus. Some of the synapses in this reflex are
electrical synapses which bypass the relatively slow process of chemical synaptic transmission by making an electrical "fusion" of the pre- and post-synaptic membranes. This adds to its rapidity.
This is considered a life preserving response for avoiding predators and has therefore been selected for rapid response times.